List of sminthids

Brown birch mouse clinging to a branch
Northern birch mouse (Sicista betulina)

Sminthidae is a family of mammals in the order Rodentia and part of the Myomorpha suborder. Members of this family are called sminthids or birch mice. They are found in Asia and Europe, primarily in forests, shrublands, and grasslands, though some species can be found in wetlands. They are all of a similar size, ranging from 4–8 cm (2–3 in) long, plus a 6–12 cm (2–5 in) tail. Sminthids are omnivores and primarily eat seeds, berries, and insects.[1] No sminthids have population estimates, but the Hungarian birch mouse is categorized as endangered, while the Armenian birch mouse, Nordmann's birch mouse, and Severtzov's birch mouse are categorized as critically endangered.

The sixteen extant species of Sminthidae are all in a single genus. Several extinct prehistoric sminthid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.[2]

Conventions

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IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically endangered (3 species)
 EN Endangered (1 species)
 VU Vulnerable (1 species)
 NT Near threatened (2 species)
 LC Least concern (9 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the sminthid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

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Sminthidae is a family consisting of sixteen extant species in a single genus, Sicista. This does not include hybrid species or extinct prehistoric species.

Sminthids

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The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]

Genus Sicista Gray, 1827 – sixteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Altai birch mouse


S. napaea
Hollister, 1912
Eastern Kazakhstan and southern Russia Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[4]

Habitat: Shrubland and grassland[5]
 LC 


Unknown Unknown[5]

Armenian birch mouse


S. armenica
Baskevich, 1988
Armenia Size: About 7 cm (3 in) long, plus 9–11 cm (4 in) tail[6]

Habitat: Grassland[7]
 CR 


Unknown Population declining[7]

Caucasian birch mouse


S. caucasica
Vinogradov, 1925
Southwestern Russia and northern Georgia Size: 4–7 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[6]

Habitat: Grassland[8]
 NT 


Unknown Unknown[8]

Chinese birch mouse

S. concolor
(Büchner, 1892)
Central and western China Size: 5–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[9]
 LC 


Unknown Unknown[9]

Gray birch mouse


S. pseudonapaea
Strautman, 1949
Eastern Kazakhstan and northwestern China Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[4]

Habitat: Forest, shrubland, and grassland[10]
 LC 


Unknown Unknown[10]

Hungarian birch mouse

S. trizona
(Frivaldszky, 1865)
Hungary and Romania Size: 5–7 cm (2–3 in) long, plus 6–10 cm (2–4 in) tail[4]

Habitat: Grassland[11]
 EN 


Unknown Population declining[11]

Kazbeg birch mouse


S. kazbegica
Sokolov, Baskevich, & Kovalskaya, 1986
Georgia and southwestern Russia Size: 6–7 cm (2–3 in) long, plus 9–11 cm (4 in) tail[6]

Habitat: Grassland[12]
 VU 


Unknown Unknown[12]

Kluchor birch mouse


S. kluchorica
Sokolov, Kovalskaya, & Baskevich, 1980
Georgia and southwestern Russia Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[6]

Habitat: Grassland[13]
 LC 


Unknown Unknown[13]

Long-tailed birch mouse


S. caudata
Thomas, 1907
Eastern Russia and eastern China Size: 6–8 cm (2–3 in) long, plus 9–12 cm (4–5 in) tail[6]

Habitat: Forest and grassland[14]
 LC 


Unknown Unknown[14]

Nordmann's birch mouse


S. loriger
(Nathusius, 1840)
Eastern Europe and southwestern Russia Size: 5–8 cm (2–3 in) long, plus 6–9 cm (2–4 in) tail[4]

Habitat: Grassland[15]
 CR 


Unknown Unknown[15]

Northern birch mouse

S. betulina
(Pallas, 1779)
Central, eastern, and northern Europe and western Asia
Map of range
Size: 5–8 cm (2–3 in) long, plus 8–10 cm (3–4 in) tail[4]

Habitat: Forest, grassland, and inland wetlands[16]
 LC 


Unknown Unknown[16]

Severtzov's birch mouse


S. severtzovi
Ogniov, 1935
Southwestern Russia
Map of range
Size: 6–8 cm (2–3 in) long, plus 7–9 cm (3–4 in) tail[4]

Habitat: Grassland[17]
 CR 


Unknown Population declining[17]

Southern birch mouse

S. subtilis
(Pallas, 1773)
Central Asia Size: 5–8 cm (2–3 in) long, plus 7–9 cm (3–4 in) tail[4]

Habitat: Shrubland and grassland[18]
 LC 


Unknown Population declining[18]

Strand's birch mouse


S. strandi
(Formozov, 1931)
Ukraine and southwestern Russia Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[4]

Habitat: Forest and grassland[19]
 LC 


Unknown Population steady[19]

Tien Shan birch mouse


S. tianshanica
(Zalensky, 1903)
Central Asia Size: 5–8 cm (2–3 in) long, plus 10–12 cm (4–5 in) tail[6]

Habitat: Forest and grassland[20]
 LC 


Unknown Unknown[20]

Tsimlyansk birch mouse


S. cimlanica
Kovalskaya, Tikhonov, Tikhonova, Surov, & Bogomolov, 2000
Ukraine and southwestern Russia Size: Unknown[4]

Habitat: Grassland[21]
 NT 


Unknown Unknown[21]

References

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  1. ^ Nowak, p. 1330
  2. ^ "Fossilworks: Dipodidae". Paleobiology Database. University of Wisconsin–Madison. Archived from the original on March 21, 2025. Retrieved September 8, 2025.
  3. ^ Wilson; Reeder, pp. 886-890
  4. ^ a b c d e f g h i Chernasky; Motis; Burgin, p. 205
  5. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Sicista napaea". IUCN Red List of Threatened Species. 2016 e.T20190A115157265. doi:10.2305/IUCN.UK.2016-3.RLTS.T20190A22204366.en.
  6. ^ a b c d e f g Chernasky; Motis; Burgin, p. 204
  7. ^ a b Rusin, M. (2020). "Sicista armenica". IUCN Red List of Threatened Species. 2020 e.T20185A22204773. doi:10.2305/IUCN.UK.2020-2.RLTS.T20185A22204773.en.
  8. ^ a b Shenbrot, G. (2020). "Sicista caucasica". IUCN Red List of Threatened Species. 2020 e.T20186A22204847. doi:10.2305/IUCN.UK.2020-2.RLTS.T20186A22204847.en.
  9. ^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Sicista concolor". IUCN Red List of Threatened Species. 2016 e.T20188A115157138. doi:10.2305/IUCN.UK.2016-3.RLTS.T20188A22204686.en.
  10. ^ a b Cserkész, T. (2025). "Sicista pseudonapaea". IUCN Red List of Threatened Species. 2025 e.T20191A279719644.
  11. ^ a b Cserkész, T. (2024). "Sicista trizona". IUCN Red List of Threatened Species. 2024 e.T92332716A221786778. doi:10.2305/IUCN.UK.2024-2.RLTS.T92332716A221786778.en.
  12. ^ a b Cserkész, T.; Gazzard, A.; Shenbrot, G. (2025). "Sicista kazbegica". IUCN Red List of Threatened Species. 2025 e.T20189A278604388.
  13. ^ a b Kennerley, R. (2017). "Sicista kluchorica". IUCN Red List of Threatened Species. 2017 e.T20196A22204093. doi:10.2305/IUCN.UK.2017-2.RLTS.T20196A22204093.en.
  14. ^ a b Cserkész, T. (2025). "Sicista caudata". IUCN Red List of Threatened Species. 2025 e.T20187A279086628.
  15. ^ a b Rusin, M. (2024). "Sicista loriger". IUCN Red List of Threatened Species. 2024 e.T110500058A221790176. doi:10.2305/IUCN.UK.2024-2.RLTS.T110500058A221790176.en.
  16. ^ a b Meinig, H.; Zagorodnyuk, I.; Henttonen, H.; Zima, J.; Coroiu, I. (2017) [errata version of 2016 assessment]. "Sicista betulina". IUCN Red List of Threatened Species. 2016 e.T20184A115156920. doi:10.2305/IUCN.UK.2016-3.RLTS.T20184A22203739.en.
  17. ^ a b Rusin, M. (2024). "Sicista severtzovi". IUCN Red List of Threatened Species. 2024 e.T221733873A221734933. doi:10.2305/IUCN.UK.2024-2.RLTS.T221733873A221734933.en.
  18. ^ a b Cserkész, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Sicista subtilis". IUCN Red List of Threatened Species. 2017 e.T91934441A123795199. doi:10.2305/IUCN.UK.2017-3.RLTS.T91934441A22204448.en.
  19. ^ a b Tsytsulina, K.; Formozov, N.; Zagorodnyuk, I.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Sicista strandi". IUCN Red List of Threatened Species. 2016 e.T20193A115157429. doi:10.2305/IUCN.UK.2016-3.RLTS.T20193A22204276.en.
  20. ^ a b Cassola, F. (2016). "Sicista tianshanica". IUCN Red List of Threatened Species. 2016 e.T20195A22204006. doi:10.2305/IUCN.UK.2016-2.RLTS.T20195A22204006.en.
  21. ^ a b Rusin, M. (2024). "Sicista cimlanica". IUCN Red List of Threatened Species. 2024 e.T221732279A221732462. doi:10.2305/IUCN.UK.2024-2.RLTS.T221732279A221732462.en.

Sources

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